I. A. Abbott and G. J. Hollenberg, Marine algae of California, 1976.

C. A. Agardh, Systema algarum. Lundae, Literis Berlingianis, 1824.

A. Bakun, B. A. Black, S. J. Bograd, A. J. Miller, R. R. Rykaczewski et al., Anticipated Effects of Climate Change on Coastal Upwelling Ecosystems, Curr. Clim. Change Rep, vol.1, pp.85-93, 2015.

R. T. Barber, Upwelling Ecosystems. Encyclopedia of Ocean Sciences: Second Edition, pp.225-232, 2008.

D. Bates, M. Maechler, B. Bolker, and S. Walker, Fitting Linear Mixed-Effects Models Using lme4, Journal of Statistical Software, vol.67, issue.1, pp.1-48, 2015.

C. A. Blanchette, Seasonal patterns of disturbance influence recruitment of the sea palm, P. palmaeformis, Journal of Experimental Marine Biology and Ecology, vol.197, issue.1, pp.1-14, 1996.

C. A. Blanchette, B. G. Miner, and S. D. Gaines, Geographic variability in form, size and survival of E. menziesii around Point Conception, California, Marine Ecology Progress Series, vol.239, pp.69-82, 2002.

J. E. Byrnes, D. C. Reed, B. J. Cardinale, K. C. Cavanaugh, S. J. Holbrook et al., Climate-driven increases in storm frequency simplify kelp forest food webs, Global Change Biology, vol.17, issue.8, pp.2513-2524, 2011.

J. H. Connell, Some mechanisms producing structure in natural communities: A model and evidence from field experiments. Ecology and evolution of communities, pp.460-490, 1975.

R. E. Corcoran, Geology of Lake Owyhee State Park and vicinity, Oregon. The Ore Bin, vol.27, pp.81-100, 1965.

P. K. Dayton, Dispersion, Dispersal, and Persistence of the Annual Intertidal Alga, P. palmaeformis Ruprecht, vol.54, pp.433-438, 1973.

P. K. Dayton, Ecology of Kelp Communities, Annual Review of Ecology and Systematics, vol.16, pp.215-245, 1985.

P. K. Dayton, M. J. Tegner, P. E. Parnell, and P. B. Edwards, Temporal and Spatial Patterns of Disturbance and Recovery in a, Kelp Forest Community. Ecological Monographs, vol.62, issue.3, pp.421-445, 1992.

D. Lorenzo, E. Schneider, N. Cobb, K. M. Franks, P. J. Chhak et al., North Pacific Gyre Oscillation links ocean climate and ecosystem change, Geophysical Research Letters, vol.35, issue.8, 2008.
URL : https://hal.archives-ouvertes.fr/hal-00634617

L. J. Falkenberg, R. , D. B. Connell, and S. D. , Stability of Strong Species Interactions Resist the Synergistic Effects of Local and Global Pollution in Kelp Forests, Plos one, vol.7, issue.3, 2012.

T. Freidenburg, G. Allison, P. Halpin, and B. Menge, Chapter 4: Effects of spatial and temporal variability in coastal upwelling on intertidal algae

S. D. Fretwell, The Regulation of Plant Communities by the Food Chains Exploiting Them, Perspectives in Biology and Medicine, vol.20, issue.2, pp.169-185, 1977.

S. D. Fretwell, Food Chain Dynamics: The Central Theory of Ecology?, Oikos, vol.50, issue.3, pp.291-301, 1987.

T. Hothorn, F. Bretz, and P. Westfall, Simultaneous Inference in General Parametric Models, Biometrical Journal, vol.50, issue.3, pp.346-363, 2008.

S. Hu and A. V. Fedorov, The extreme El Niño of 2015-2016 and the end of global warming hiatus, Geophysical Research Letters, vol.44, issue.8, pp.3816-3824, 2017.

M. G. Jacox, E. L. Hazen, K. D. Zaba, D. L. Rudnick, C. A. Edwards et al., Impacts of the 2015-2016 El Niño on the California Current System: Early assessment and comparison to past events, Geophysical Research Letters, vol.43, issue.13, 2016.

A. Kassambara and F. Mundt, Package 'factoextra'. Extract and Visualize the Results of Multivariate Data Analyses, 2017.

M. T. Kavanaugh and B. A. Menge, Phytoplankton Shading of Marine Benthic Macrophytes: Implications for Intertidal Community Structure, pp.74-77, 2005.

S. Korpinen, V. Jormalainen, and T. Honkanen, Effects of nutrients, herbivory, and depth on the macroalgal community in the rocky sublittoral, Ecology, vol.88, issue.4, pp.839-852, 2007.

K. A. Krumhansl, D. K. Okamoto, A. Rassweiler, M. Novak, J. J. Bolton et al., Global patterns of kelp forest change over the past half-century, Proceedings of the National Academy of Sciences, vol.113, issue.48, pp.13785-13790, 2016.

D. Mchugh, A guide to the seaweed industry. FAO Fisheries Technical Paper, vol.441, p.105, 2003.

B. A. Menge, Top-down and bottom-up community regulation in marine rocky intertidal habitats, Journal of Experimental Marine Biology and Ecology, vol.250, issue.1-2, pp.257-289, 2000.

B. A. Menge and G. M. Branch, Rocky intertidal communities. Marine community ecology, pp.221-246, 2001.

B. A. Menge, F. Chan, K. J. Nielsen, E. Lorenzo, and J. Di-&-lubchenco, Climatic variation alters supply-side ecology: impact of climate patterns on phytoplankton and mussel recruitment, Ecological Monographs, vol.79, issue.3, 2009.

B. A. Menge, B. Daley, and P. A. Wheeler, Control of Interaction Strength in Marine Benthic Communities, 1996.

B. A. Menge, T. C. Gouhier, S. D. Hacker, F. Chan, and K. J. Nielsen, Are metaecosystems organized hierarchically? A model and test in rocky intertidal habitats, Ecological Monographs, vol.85, issue.2, pp.213-233, 2015.

B. A. Menge and A. M. Olson, Role of scale and environmental factors in regulation of community structure, Trends Ecol Evol, vol.5, issue.2, p.90048, 1990.

B. A. Menge and J. P. Sutherland, Community Regulation: Variation in Disturbance, Competition, and Predation in Relation to Environmental Stress and Recruitment, The American Naturalist, vol.130, pp.730-757, 1987.

B. A. Menge and J. P. Sutherland, Species Diversity Gradients: Synthesis of the Roles of Predation, Competition, and Temporal Heterogeneity, The American Naturalist, vol.110, pp.351-369, 1976.

L. Oksanen, S. D. Fretwell, J. Arruda, and P. Niemela, Exploitation Ecosystems in Gradients of Primary Productivity, The American Naturalist, vol.118, issue.2, pp.240-261, 1981.

R. T. Paine, Disaster, Catastrophe, and Local Persistence of the Sea Palm P. palmaeformis, Advancement Of Science, issue.4407, pp.685-687, 1979.

R. T. Paine, Habitat Suitability and Local Population Persistence of the Sea Palm P. palmaeformis, Ecology, vol.69, issue.6, pp.1787-1794, 1988.

D. Roemmich and J. Mcgowan, American Association for the Advancement of Science, vol.217, pp.1161-1163, 2014.

B. D. Russell and S. D. Connell, A novel interaction between nutrients and grazers alters relative dominance of marine habitats, Marine Ecology Progress Series, vol.289, pp.5-11, 2005.

W. J. Sydeman, D. S. Schoeman, R. R. Rykaczewski, S. A. Thompson, B. A. Black et al., Climate change and wind intensification in coastal upwelling ecosystems, Science, vol.345, issue.6192, pp.77-81, 2014.

L. R. Vinueza, G. M. Branch, M. L. Branch, and R. H. Bustamante, Top-down Herbivory and Bottom-up El Niño Effects on Galápagos Rocky-Shore Communities, Ecological Monographs, vol.76, issue.1, pp.111-131, 2006.

D. Wang, T. C. Gouhier, B. A. Menge, and A. R. Ganguly, Intensification and spatial homogenization of coastal upwelling under climate change, Nature, vol.518, issue.7539, pp.390-394, 2015.

T. Wernberg, M. S. Thomsen, F. Tuya, G. A. Kendrick, A. Peter et al., Decreasing resilience of kelp beds along a latitudinal temperature gradient: potential implications for a warmer future, Ecology Letters, vol.13, pp.685-694, 2010.

C. C. Wilmers, J. A. Estes, M. Edwards, K. L. Laidre, and B. Konar, Do trophic cascades affect the storage and flux of atmospheric carbon? An analysis of sea otters and kelp forests, Frontiers in Ecology and the Environment, vol.10, issue.8, pp.409-415, 2012.

P. Xiu, F. Chai, E. N. Curchitser, and F. S. Castruccio, Future changes in coastal upwelling ecosystems with global warming: The case of the California Current System, Scientific Reports, vol.8, issue.1, pp.1-9, 2018.

, 18) National centers for environmental information, NOAA-National Oceanic and Atmospheric Administration URL, NOAA-National Oceanic and Atmospheric Administration URL

, NOAA-National Oceanic and Atmospheric Administration URL

, CB-2018, vol.9466, p.2016, 2016.

, SiteFC

, SiteSH

2. Table, Pairwise comparison in length post-hoc results from experimental analysis using Tukey for each species. Species Estimate Std. Error Z ratio p.value Saccharina Sessilis, p.2016, 2016.

, CF-2016,CP-7.6922895 1.0210607-11.047 P<0, CF-2018, vol.001, p.2017, 2016.