A. Bio and /. Oc, , 2017.

. Agreste, Mémento de la statistique agricole. La documentation française, p.44, 2015.

. Agreste, Agreste Conjecture : Viticulture, juillet, vol.2019, pp.2019-106, 2019.

, Site du ministère de l'agriculture et de l'alimentation, 2018.

M. Angaud, Caractérisation de la flore adventice en viticulture : Importance du paysage, du pédo-climat et des pratiques agricoles dans la structuration des communautés, Stage 4 ème année, UniLasalle Rouen, vol.18, 2018.

L. Armengot, J. M. Blanco-moreno, P. Bàrberi, G. Bocci, S. Carlesi et al., Tillage as a driver of change in weed communities: a functional perspective, Agriculture, Ecosystems and Environment, vol.222, pp.276-285, 2016.

H. G. Baker, The evolution of weeds, Annual review of Ecology and Systematics, vol.5, pp.1-24, 1974.

P. Bàrberi, G. Burgio, G. Dinelli, A. C. Moonen, S. Otto et al., Functional biodiversity in the agricultural landscape: relationships between weeds and arthropod fauna, Weed Research, vol.50, pp.388-401, 2010.

G. Barralis, G. Cloquemin, and A. Et-guerin, Evolution de la flore adventice du vignoble de Côte-d'Or sous la pression des techniques d'entretien des cultures, Agronomie, EDP Sciences, vol.3, pp.585-594, 1983.

K. Baumgartner, K. L. Steenwerth, and L. Veilleux, Cover-crop systems affect weed communities in a California vineyard, Weed Science, vol.56, pp.596-605, 2008.

H. Bestovà, F. Munoz, P. Svoboda, P. Skaloud, and C. Violle, Ecological and biogeographical drivers of freshwater green algae biodiversity: from local communities to large-scale species pools of desmids, Oecologia, vol.186, pp.1017-1030, 2018.

J. Bennie, M. O. Hill, R. Baxter, and B. Huntley, Influence of slope and aspect on long-term vegetation change in British chalk grasslands, Journal of Ecology, vol.94, pp.355-368, 2006.

T. Boisson, Influence du paysage sur la flore adventice en viticulture, Stage M2 sciences du territoire : université Grenoble Alpes, vol.53, 2016.
URL : https://hal.archives-ouvertes.fr/dumas-01442208

D. Borcard, F. Gillet, and P. Legendre, Numerical ecology with R, vol.306, pp.978-979, 2011.
URL : https://hal.archives-ouvertes.fr/hal-00530195

Z. Botta-dukát, Rao's quadratic entropy as a measure of functional diversity based on multiple traits, Journal of Vegetation Science, vol.16, pp.533-540, 2005.

B. Bourgeois, F. Minoz, G. Fried, L. Mahaut, L. Armengot et al., What makes a weed a weed? A large scale evaluation of arable weeds through a functional lens, American Journal of Botany, vol.106, pp.1-11, 2019.
URL : https://hal.archives-ouvertes.fr/hal-02009206

O. T. Bruggisser, M. H. Schmidt-entling, and S. Bacher, Effects of vineyard management on biodiversity at three trophic levels, Biological Conservation, vol.143, pp.1521-1528, 2010.

L. A. Brun, L. Corff, J. Maillet, and J. , Effects of elevated soil copper on phenology, growth and reproduction of five ruderal plant species, Environmental Pollution, vol.122, pp.312-315, 2003.

L. Cazenave, Flore adventice des vignes en Gironde, Comité de pilotage : Chambre d'Agriculture de la Gironde, vol.69, 2017.

F. Celette, A. Ripoche, and C. Gary, WaLIS_ A simple model to simulate water partitioning in a crop association: The example of an intercropped vineyard, Agriculture Water Management, vol.97, pp.1749-1759, 2010.

B. Chauvel, H. Darmency, N. Munier-jolain, and A. Et-rodriguez, Gestion durable de la flore adventice des cultures, Editions Quae, vol.354, pp.978-980, 2018.

M. Chytrý, L. Tichý, J. Holt, and Z. Botta-dukát, Determination of diagnostic species with statistical fidelity measures, Journal of Vegetation Science, vol.13, pp.79-90, 2002.

W. K. Cornwell, D. W. Schwilk, and D. D. Ackerly, A trait-based test for habitat filtering: convex hull volume, Ecology, vol.87, pp.1465-1471, 2006.

S. M. Dabney, J. A. Delgado, and D. W. Reeves, Using winter cover crops to improve soil and water quality, Communication in Soil Science and Plant Analysis, vol.32, pp.1221-1250, 2001.

R. Dion, . Histoire-de-la-vigne, and F. Du-vin-en, , vol.768, pp.2-08, 1977.

S. Díaz and M. Cabido, Vive la difference: Plant functional diversity matters to ecosystem processes, Trends in Ecology and Evolution, vol.16, issue.01, pp.2283-2285, 2001.

S. Dray, P. Choler, S. Dolédec, P. R. Peres-neto, W. Thuiller et al., Combining the fourth-corner and the RLQ methods for assessing trait responses to environmental variation, Ecology, vol.95, pp.14-21, 2014.
URL : https://hal.archives-ouvertes.fr/halsde-00942845

H. Ellenberg, H. E. Weber, R. Düll, V. Wirth, W. Werner et al., Zeigerwerte von Pflanzen in MittelEuropa, Scripta Geobotanica, vol.18, pp.1-258, 1992.

H. Ellenberg, Vegetation Mitteleuropas mit den Alpen in ökologischer, dynamischer und historischer Sicht, pp.3-8001, 1095.

G. Fried, B. Chauvel, and X. Reboud, A functional analysis of large-scale temporal shifts from 1970 to 2000 in weed assemblages of sunflower crops in France, Journal of Vegetation Science, vol.20, pp.49-58, 2009.

G. Fried, E. Kazakou, and S. Gaba, Trajectories of weed communities explained by traits associated with species' response to management practices. Agriculture, Ecosystems and Environment, vol.158, pp.147-155, 2012.
URL : https://hal.archives-ouvertes.fr/hal-01506122

G. Fried, F. Dessaint, and X. Reboud, Local and regional changes in taxonomic and functional diversity of arable weed communities in Burgundy (France) between 1970s and the, Botany Letters, vol.163, pp.359-371, 2000.
URL : https://hal.archives-ouvertes.fr/hal-01602252

G. Fried and J. Maillet, Chapitre 2 : Diversité et réponses de la flore des champs cultivés à l'évolution des pratiques agricoles en France, Editions Quae, vol.354, pp.978-980, 2018.

G. Fried, A. Villiers, and E. Porcher, Assessing non-intended effects of farming practices on field margin vegetation with a functional approach, Agriculture, Ecosystems and Environment, vol.261, pp.33-44, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01763788

G. Fried, S. Cordeau, A. Metay, and E. Kazakou, Relative importance of environmental factors and farming practices in shaping weed communities structure and composition in French vineyards, Agriculture, Ecosystems and Environment, vol.275, pp.1-13, 2019.

G. Fried, Apports des approches fonctionnelles pour l'évaluation des risques associés aux changements de végétation induits par les activités humaines, Mémoire pour l'obtention de l'habilitation à diriger des recherches : Amphithéâtre de la délégation régionale du, vol.142, 2019.

R. J. Froud-williams, D. S. Drennan, and R. J. Chancellor, Influence of cultivation regime on weed floras of arable cropping systems, Journal of Applied Ecology, vol.20, pp.187-197, 1983.

S. Gaba, G. Fried, E. Kazakou, B. Chauvel, and M. L. Navas, Agroecological weed control using a functional approach: a review of cropping systems diversity. Agronomy for Sustainable Development, vol.34, pp.103-119, 2014.
URL : https://hal.archives-ouvertes.fr/hal-01234789

S. Gaba, R. Perronne, G. Fried, A. Gardarin, F. Bretagnolle et al., Response and effect traits of arable weeds in agro-ecosystems: a review of current knowledge, Weed research, vol.57, pp.123-147, 2017.
URL : https://hal.archives-ouvertes.fr/hal-01547815

P. Gago, C. Cabaleiro, and J. Garcia, Preliminary study of the effect of soil management systems on the adventitious flora of a vineyard in northwestern Spain, Crop protection, vol.26, pp.584-591, 2007.

L. Garcia, F. Celette, C. Gary, A. Ripoche, H. Valès-gomez et al., Management of service crops for the provision of ecosystem services in vineyards: A review. Agriculture, Ecosystems and Environment, vol.251, pp.158-170, 2018.
URL : https://hal.archives-ouvertes.fr/hal-01614417

E. Garnier, M. L. Navas, and K. Grigulis, Plant functional diversity organism traits, community structure and ecosystem properties, vol.256, pp.978-0198757375, 2016.

J. Gasquez, B. Chauvel, H. Darmency, N. Munier-jolain, and A. Et-rodriguez, Chapitre 1 : Historique de la gestion de la flore adventice, Editions Quae, vol.354, pp.978-980, 2018.

C. Gaviglio, La gestion des sols viticoles, IFV -La France Agricole, Vigne et vin, p.9782855572581, 2013.

J. P. Grime, Vegetation classification by reference to strategies, Nature, vol.250, pp.26-31, 1974.

J. P. Grime, Plant strategies and vegetation processes, Journal of Ecology, vol.68, pp.704-706, 1979.

J. P. Grime, The CSR model of primary plant strategies-origins, implications and tests, Plant Evolutionary Biology, 1988.

J. P. Grime, Plant Strategies, Vegetation Processes and Ecosystem Properties, vol.417, pp.0-471, 2001.

J. P. Grime, Trait convergence and trait divergence in herbaceous plant communities: mechanisms and consequences, Journal of Vegetation Science, vol.17, pp.255-260, 2006.

J. P. Grime and S. Pierce, The evolutionary strategies that shape ecosystems, 2012.

R. M. Gunton, S. Petit, and S. Gaba, Functional traits relating arable weed communities to crop characteristics, Journal of Vegetation Science, vol.22, pp.541-550, 2011.

D. A. Heemsbergen, M. P. Berg, M. Loreau, J. R. Van-hal, J. H. Faber et al., Biodiversity effects on soil processes explained by interspecific functional dissimilarity, Science, vol.306, pp.1019-1020, 2004.

M. O. Hill and A. J. Smith, Principal component analysis of taxonomic data with multi-state discrete characters, Taxon, vol.25, pp.249-255, 1976.

M. O. Hill, J. O. Mountford, D. B. Roy, and R. G. Bruce, Ellenberg's indicator values for British plants. ECOFACT Volume 2 Technical Annex, vol.46, p.1870393, 1999.

J. G. Hodgson, P. J. Wilson, R. Hunt, J. P. Grime, and K. Thompson, Allocating CSR plant functional types: a soft approach to a hard problem, Oikos, vol.85, pp.282-294, 1999.

D. G. Hole, A. J. Perkins, J. D. Wilson, I. H. Alexander, P. V. Grice et al., Does organic farming benefit biodiversity? Biological Conservation, vol.122, pp.113-130, 2005.

F. D. Hulot, G. Lacroix, F. O. Lescher-moutoue, and M. Loreau, Functional diversity governs ecosystem response to nutrient enrichment, Nature, vol.405, pp.340-344, 2000.

C. M. Hulshof, C. Violle, M. J. Spasojevic, B. Mcgill, E. Damschen et al., Intra-specific and inter-specific variation in specific leaf area reveal the importance of abiotic and biotic drivers of species diversity across elevation and latitude, Journal of Vegetation Science, vol.24, pp.921-931, 2013.

F. Husson, S. Lê, and J. Et-pagès, Analyses de données avec R. Presses universitaires de Rennes, 2 ème édition, vol.240, pp.978-980, 2016.

. Ivf-occitanie, Institut français de la vigne et du vin, Occitanie. Consultable, 2019.

P. Julve, Baseflor: Index botanique, écologique et chronologique de la flore de France, 1998.

E. K. Karadimou, A. S. Kallimanis, I. Tsiripidis, and P. Dimopoulos, Functional diversity exhibits a diverse relationship with area, even a decreasing one, Scientific Reports, vol.6, p.35420, 2016.

E. Kazakou, G. Fried, J. Richarte, O. Gimenez, C. Violle et al., A plant trait-based response-and-effect framework to assess vineyard inter-row soil management, Botany letters, vol.163, pp.373-388, 2016.

E. Kazakou, F. Jacquemin, M. L. Navas, and C. Violle, Chapitre 5 : Fonctions et services des plantes adventices dans les agroécosystèmes in Chauvel B., Darmency H., Munier-Jolain N. et Rodriguez A. Gestion durable de la flore adventice des cultures, Editions Quae, vol.354, pp.978-980, 2018.

M. Kleyer, R. M. Bekker, I. C. Knevel, J. P. Bakker, K. Thompson et al., The LEDA Traitbase: A database of life-history traits of North West European flora, Journal of Ecology, vol.96, pp.1266-1274, 2008.

S. Klotz, I. Kühn, and D. W. , BIOLFLOR -Eine Datenbank zu biologisch-ökologischen Merkmalen der Gefä?pflanzen in Deutschland, Schriftenreihe für Vegetationskunde, 38. Bundesamt für Naturschutz, vol.333, 2002.

S. Kullback and R. A. Leibler, On information and sufficiency, The Annals of Mathematical Statistics, vol.22, pp.79-86, 1951.

E. Laliberté and P. Legendre, A distance-based framework for measuring functional diversity from multiple traits, Ecology, vol.91, pp.299-305, 2010.

J. Leps, F. De-bello, S. Lavorel, and S. Berman, Quantifying and interpreting functional diversity of natural communities: practical considerations matter, Preslia, vol.78, pp.481-501, 2006.
URL : https://hal.archives-ouvertes.fr/halsde-00293183

X. Louchart, M. Voltz, P. Andrieux, and R. Moussa, Herbicide transport to surface waters at field and watershed scales in a Mediterranean vineyard area, Journal of Environmental Quality, vol.30, pp.982-991, 2001.

Z. Lososovà, J. Danihelka, and M. Chytrý, Seasonal dynamics and diversity of weed vegetation in tilled and mulched vineyards, Biologia, vol.58, pp.49-57, 2003.

C. Maclaren, J. Bennett, and K. Dehnen-schmutz, Management practices influence the competitive potential of weed communities and their value to biodiversity in South African vineyards, Weed Research, vol.59, pp.93-106, 2019.

J. Maillet, Constitution et dynamique des communautés de mauvaises herbes des vignes de France et des rizières de Camargue, vol.179, 1992.

R. C. Mart??ez, E. R. Gonzalo, M. E. Laespada, S. Roman, and F. J. , Evaluation of surface-and ground-water pollution due to herbicides in agricultural areas of Zamora and Salamanca (Spain), Journal of Chromatography A, vol.869, pp.1188-1195, 2000.

N. W. Mason, F. De-bello, D. Mouillot, S. Pavoine, and S. Dray, A guide for using functional diversity indices to reveal changes in assembly processes along ecological gradients, Journal of Vegetation Science, vol.24, pp.794-806, 2013.
URL : https://hal.archives-ouvertes.fr/hal-02281571

H. Meiss, S. Médiène, and R. Waldhardt, Perennial Lucerne affects weed community trajectories in grain crop rotations, Weed Research, vol.50, 2010.
URL : https://hal.archives-ouvertes.fr/hal-01644347

A. Metay, L. Garcia, and C. Gary, Chapitre 15 : Gestion durable de la flore adventice en culture pérenne : enjeux et stratégies en viticulture, Editions Quae, vol.354, pp.978-980, 2018.

D. Mezière, C. Gary, J. M. Barbier, P. Rio, L. Bernos et al., Ecophyto R&D, vers des systèmes de cultures économes en produits sanitaires. Tome III, analyses comparatives de différents systèmes en viticulture, vol.57, 2009.

, Millenium Ecosystem Assessment. Ecosystems and human well-being: synthesis world resources institute, vol.155, pp.1-59726, 2005.

J. Montégut, Pérennes et vivaces nuisibles en Agriculture. Ed. sous l'égide de Monsanto, J. Manuel Ed. Insenvelop, vol.414, 1982.

D. Moreau, G. Milard, and N. Munier-jolain, A plant nitrophily index based on plant leaf area response to soil nitrogen availability. Agronomy for Sustainable Development, vol.33, pp.809-815, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01201394

J. Nascimbene, L. Marini, and M. G. Paoletti, Organic farming benefits local plant diversity in vineyard farms located in intensive agricultural landscape, Environmental Management, vol.49, pp.1054-1060, 2012.

M. L. Navas and E. Et-garnier, Diversité fonctionnelle des plantes : traits des organismes, structure des communautés, propriétés des écosystèmes. Editions De Boeck, vol.353, pp.978-980, 2013.

A. Pellegrino, E. Lebon, T. Simonneau, and J. Wery, Towards a simple indicator of water stress in grapevine (Vitis vinifera L.) based on the differential sensitivities of vegetative growth components, Australian Journal of Grape and Wine Research, vol.11, pp.306-315, 2005.

N. Pérez-harguindeguy, S. Diaz, E. Garnier, S. Lavorel, H. Poorter et al.,

W. K. Cornwell, J. M. Craine, D. E. Gurvich, C. Urcelay, E. J. Veneklaas et al., New handbook for standardised measurement of plant functional traits worldwide, Australian Journal of Botany, vol.61, pp.167-234, 2013.

F. Perring, Topography gradients of chalk grassland, Journal of Ecology, vol.47, pp.447-481, 1959.

R. Phillips and . French, Wine: A history, vol.335, p.9780520285231, 2016.

S. Pierce, D. Negreiros, B. E. Cerabolini, J. Kattge, S. Díaz et al., A global method for calculating plant CSR ecological strategies applied across biomes world-wide, Functional Ecology, vol.31, pp.444-457, 2017.

E. Plan and . Ii, Ministère de l'agriculture, de l'agroalimentaire et de la forêt et le ministère de l'Ecologie, du développement durable et de l'énergie. 67 pages, 2015.

L. Porcova and J. Wikler, The effect of the different vineyards management to composition of weed species, Conference Mentel Net, pp.90-94, 2014.

D. Posada and T. R. Buckley, Model selection and model averaging in Phylogenetics: Advantages of Akaike Information Criterion and Bayesian approaches over likelihood ratio tests, Systematic biology, vol.53, pp.793-808, 2004.

S. R. Radosevich, J. S. Holt, and C. M. Ghersa, Ecology of weeds and invasive plants: relationships to agriculture and natural resources management, vol.454, 2007.

J. M. Randall, (eds) Assessment and management of plant invasions, pp.978-979, 1997.

I. Roschewitz, D. Gabriel, T. Tscharntke, and C. Thies, The effect of landscape complexity on arable weed species diversity in organic and conventional farming, Journal of Applied Ecology in press, vol.42, pp.873-882, 2005.

, Seed Information Database (SID), Royal Botanic Gardens Kew, 2019.

M. Ruiz-colmenero, R. Bienes, D. Eldridge, and M. Marques, Vegetation cover reduces erosion and enhances soil organic carbon in a vineyard in the central Spain, Catena, vol.104, pp.153-160, 2013.

A. Saatkamp, L. Affre, P. Poschlod, P. Roche, U. Deuil et al., Decrease of plant alpha and beta-diversity with management intensity in vineyards and the influence of landscape context, Ecologia Mediterranea, vol.40, pp.17-28, 2014.

D. Schleuter, M. Daufresne, F. Massol, and C. Argillier, A user's guide to functional diversity indices, vol.80, pp.469-484, 2010.

A. Shrestha and M. Fidelibus, Grapevine row orientation affects light environment, growth, and development of black nightshade (Solanum nigrum), Weed Science, vol.53, pp.802-812, 2005.

S. Tautehahn, H. Heilmeier, L. Götzenberger, S. Koltz, C. Wirth et al., On the biogeography of seed mass in Germany -distribution patterns and environmental correlates, Ecography, vol.31, pp.457-468, 2008.

J. P. Theau, P. Cruz, D. Fallour, C. Jouany, E. Lecloux et al., Une méthode simplifiée de relevé botanique pour une caractérisation agronomique des prairies permanentes, Fourrages, vol.201, pp.19-25, 2010.

D. Tilman, J. Knops, D. Wedin, P. Reich, M. Ritchie et al., The influence of functional diversity and composition on ecosystem processes, Science, vol.277, pp.1300-1302, 1997.

J. M. Tison, et Foucauld B (coords). Flora Gallica, p.2366620128, 1196.

A. Trichard, A. Alignier, B. Chauvel, and S. Petit, Identification of weed community traits response to conservation agriculture, Agriculture, Ecosystems and Environment, vol.179, pp.179-186, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01210112

C. Van-leeuwen and G. Seguin, Incidences de l'alimentation en eau de la vigne, appréciée par l'état hydrique du feuillage, sur le développement de l'appareil végétatif et la maturation du raisin (Vitis vinifera variété Cabernet franc, Journal International des Sciences de la Vigne et du Vin, vol.28, pp.81-110, 1990.

A. M. Varela, Conception de méthodes simplifiées de relevés floristiques en viticulture à destination des réseaux DEPHY/FERME et EXPE. Mémoire en vue de l'obtention du master FACE, vol.66, 2015.

. Vin-&-vigne, Guide des vins et des vignes de France, 2015.

C. Violle, M. Navas, D. Vile, E. Kazakou, C. Fortunel et al., Let the concept of trait be functional! Oikos, vol.116, pp.882-892, 2007.

S. Vrsic, A. Ivancic, B. Pulko, and J. Valhuber, Effect of soil management systems on erosion and nutrition loss in vineyard on steep slopes, Journal of Environmental Biology, vol.32, pp.289-294, 2011.

P. S. White and S. T. Picklett, Natural disturbance and patch dynamics, an introduction, pp.3-13, 1985.

, Enseignantes référentes : Guénola Pérès et Edith Le cadre

, Auteur(s) : Cécile Blanchet Date de naissance* : 20, vol.01, 1995.

, Annexe(s): 8

A. De-soutenance, , 2019.

, Organisme d'accueil : LSV -Laboratoire de Santé des Plantes -ANSES 755 Avenue du Campus Agropolis CS 30016 34988 Montferrier-sur-lez Cedex Maîtres de stage : Guillaume Fried, Stéphane Cordeau Titre français : Les traits de réponse de la flore adventices permettent-ils de comprendre l

, Titre anglais: Can the traits help us understanding the response of weed flora to viticultural practices?